Abstract ---The validity of the Late Visean genus  Lusitanoceras Pereiro de Sousa, 1923 (Goniatitidae) is supported, and its taxonomy and stratigraphic distribution are discussed. The genus  Lusitanoceras, which has a diachronous distribution in the Uralian-Kazakhstanian-Central Asian Region on the one hand and in the Subvariscan Realm on the other, possibly evolved from  Arnsbergites. The study of the early whorls of  Lusitanoceras and  Dombarites does not support the hypothesis of the evolution of  Dombarites from  Lusitanoceras. The analysis of the stratigraphic distribution and shell morphology of these genera suggests that the genus  Dombarites evolved directly from the genus  Goniatites, while the lineage  Arnsbergites  [arrow] Lusitanoceras was most likely an evolutionary offshoot of the family Goniatitidae. The analysis of the taphonomy and lithology of  Lusitanoceras occurrences shows that in the Uralian-Kazakhstanian-Central Asian Region that this genus inhabited territories around the outer margin of carbonate platforms in an environment of intensive carbonate sedimentation. These habitats are very much different from the habitats of  Lusitanoceras in the Subvariscan Realm.