Locality Layer
China, Guangxi, Nandan region unrecognized
France, Bretagne, Kerdréolet Kerdréolet Fm.
Germany, Harz, Grumbacher Teich Goslar Shales
Germany, Rhenish Mountains, Rupbachtal, Langscheidt Mine Wissenbach Shales
Morocco, Tafilalt, Gara Mdouara E unit Klug 2001
Morocco, Tafilalt, Hamar Laghdad Upper Eifelian
Spain, Palencia, Arbejal Gustalapiedra Fm.
Spain, Palencia, Arroyo de Man Gustalapiedra Fm.
USA, Alaska Cheeneetnuk Lst. 82 m (79RB9)